How to calculate shannon wiener diversity index formula. Shannon wiener index is defined and given by the following function. H has its foundations in information theory and represents the uncertainty about the. The shannon wiener diversity index is one measure that we will use to try to draw information from samples in the field. Pdf modification of shannonwiener diversity index towards. The shannonwiener diversity index, h, is calculated using the following equation. In the shannon index, p is the proportion nn of individuals of one particular species found n divided by the total number of individuals found n, ln is the natural log. Shannons diversity index,shannonwienershannonweaver s. Posted on 201021 20170904 author bpmsg categories online tools, other articles tags berger parker. A simple, handly and powerful tool, called the dimo model based on the shannon wiener index, has been developed for analysis and evaluation of species diversity at plant community alpha level.
Posted on 201021 20170904 author bpmsg categories online tools, other articles tags berger parker, calculation, excel, shannon entropy, shannon weaver index, shannon wiener index, simpson index. If the e value is 1, the species are equally present in the habitat. Equitability assumes a value between 0 and 1 with 1 being complete evenness. Like simpsons index, shannon s index accounts for both abundance and evenness of the species present. The formula for calculating the value o f the index is. Calculate the shannon diversity index and evenness for these sample values. This is the simplest of all the measures of species diversity. Pdf a simple formula for calculating the lower limit of.
The shannon index has been a popular diversity index in the ecological literature tandon et al. The degree of uncertainty of predicting the species of a random sample is related to the diversity of a community. A simple, handly and powerful tool, called the dimo model based on the shannonwiener index, has been developed for analysis and evaluation of species diversity at plant community alpha level. A measure used by ecologists when a system contains too many individuals for each to be identified and examined. This step involved the calculation of diversity indices and evenness of.
The maximum diversity hmax of a sample is found when all species are equally abundant. This makes the shannon index and the simpson diversity or gini coefficient special cases of a more general index. I am in need of calculating shannons diversity index of my identified diatom species using excel with the formula h. Pdf a simple formula for calculating the lower limit of shannons. Simpsons diversity indexis used to calculate a measure of diversity, taking into account the. On some occasions it is called the shannonwiener index and on other occasions it is called the shannonweaver index. This diversity measure is based on information theory. Pdf the purpose of this note is to demonstrate that the exact value of the lower limit of shannons diversity index. First, enter the number of species, and then enter the name you wish to give the species, if available, and the given populations for each of the species. The diversity index was calculated by using the shannon wiener diversity index 1949. Shannon index is a commonly used diversity index that takes into account.
Shannon diversity is very widely used index for comparing diversity between various habitats clarke and warwick, 2001. Demonstration of how to calculate the shannonwiener diversity index in microsoft excel. A greater number of species and a more even distribution both increase diversity as measured by h. Nov 27, 2015 an example of the shannon wiener diversity index for science students. Relative to other diversity indices, such as simpsons index, it is considered sensitive to the addition of rare species krebs, 1989. H pilnpi where pi is the proportion of each species in the sample. Simpson diversity and the shannonwiener index as special. It combines both evenness and richness in a single measure.
Introduction, simpsons index and shannonweiner index. Dpsi log pi, where s is the total number of species in the sample, i is the total number of individuals in. Posted on 201021 20170904 author bpmsg categories online tools, other articles tags berger parker, calculation, excel, shannon entropy, shannon weaver index, shannonwiener index, simpson index. It was calculated in order to know the species diversity in different habitat hutchison, 1970 based on the abundance of the species by the following formula. On some occasions it is called the shannon wiener index and on other occasions it is called the shannon weaver index. Here p i is the proportion of total number of species made up of the ith. Application of shannons index to study diversity with. Shannonweiner index the shannon index is affected by both the number of species and their equitability, or evenness. Price, 1975, where it is known as shannon s diversity index, the shannon wiener index, the shannon weaver index and the shannon entropy.
Like simpsons index, shannons index accounts for both abundance and evenness of the species present. A diversity index also called phylogenetic or simpsons diversity index is a quantitative measure that reflects how many different types such as species there are in a dataset a community and that can simultaneously take into account the phylogenetic relations among the individuals distributed among those types, such as richness, divergence or evenness. In a low diversity brackish water habitat the diversity of the copepod community is best measured with the shannonwiener information function and its evenness by an index proposed by the first author eh181. The shannon diversity index h is another index that is commonly used to characterize species diversity in a community. Choose this diversity calculation from the alpha diversity tab. The most common measures of biodiversity are species richness, simpsons index and shannonwiener index. Species diversity concepts describing communities species.
Find shannons index of diversity and index of relative diversity for a random sample of 25 observations distributed among five categories as shown in range b4. Divide shannons diversity index h by natural logarithm of species richness lns to calculate the species evenness. I am not an ecologist and i dont know the shannonwiener index and dont remember by heart the definition of the simpsons index. Shannon wiener diversity index 18 i i s i i s i h p p p p i 2 1 1 ln log 3. A diversity index is the measure of species diversity in a given community. Species diversity and richness 4 is designed for professional ecologists and their students. Diversity of classes shannon, simpson, species statsdirect. This tutorial explains how to calculate the shannon wiener diversity index and evenness. Many indices for measuring species diversity have been proposed.
Species evenness, richness, and diversity indices as shannon weiner shannon and weaver, 1949 and simpson index simpson, 1949 were used to evaluate the bird species diversity. Note that species evenness ranges from zero to one, with zero signifying no evenness and one, a complete evenness. This was shown by comparing the statistical behaviour of the more important diversity and evenness indices currently used. It is different from species richness in that unlike richness it also shows community composition and takes into account the relative abundance of species that are present in the community. Another measure of homogeneity is given by the formula.
It turns out that the mathematical relationships hold true whether one is dealing with molecules in solution or species in an ecological community. We suggest an explanation for this dual use of terms and in so doing we offer a tribute to the late claude shannon who passed away on 24 february 2001. Putting the values into the formula for shannon index, h s 1. They are just indices of diversity, in the same way that the diameter of a sphere is an index of its volume but is not itself the volume. Since by definition the pis will all be between zero and one, the. In this article, a link is noted between a common family of diversity indices and non. A diversity index is a mathematical measure of species diversity in a community. Diversity indices like the shannon entropy shannonwiener index and the ginisimpson index are not themselves diversities.
However, if you were to include the definitions of those indices, i and others might be able to answer without having to search the literature for those definitions. A tribute to claude shannon 19162001 and a plea for more rigorous use of species richness, species diversity and the shannon wiener index. P i fraction of the entire population madeup of species i proportion of a species i relative to total number of. The most common measures of biodiversity are species richness, simpsons index and shannon wiener index. Biodiversity calculator this calculator is free to use and is designed for biologists, ecologists, teachers, and students needing to quickly calculate the biodiversity indexes of an ecosystem. The calculator uses the following formula to calculate the shannonwiener diversity index. There have been numerous attempts to create compound indices that combine measures of richness and abundance. Shannons diversity index real statistics using excel.
Natural resource biometrics measures of diversity shannon. Foremost among these are the shannons diversity h and simpsons diversity d 1 indices table table1, 1, which differ in their theoretical foundation and interpretation magurran 2004. Study of shrub and grassy herbal diversity in naturalstand. Shannonwiener index is defined and given by the following function. Biodiversity calculator for the simpson and shannon indexes.
In fact, in the late 1940s shannon had built on the work of wiener. Shannonwiener diversity index 18 i i s i i s i h p p p p i 2 1 1 ln log 3. Shannon weiner index assumes that individuals are randomly sampled from an independent large population and all the. Jan 26, 2012 shannon index is a commonly used diversity index that takes into account both abundance and evenness of species present in the community. Interestingly shannon, a physicist, developed the index as a formula for measuring the entropy of matter in the universe. A simple formula for calculating the lower limit of shannons diversity index. Jan 06, 2011 demonstration of how to calculate the shannon wiener diversity index in microsoft excel. Measuring diversity has been of historical significance and due to the obvious declines in habitat diversity this data is still valuable. A tribute to claude shannon 19162001 and a plea for. N2 varies from one to s the number of species in the sample. How can i calculate shannons diversity index using excel sheet. Shannonwiener diversity function h pi lnpi h shannonwiener index of species diversity s number of species in community pi proportion of total abundance represented by ith species species diversity 45. Diversity indices provide more information about community composition than simply species richness i. Pi in pi where pi s n s number of individuals of one species n total number of all individuals in the sample in logarithm to base e measurement of species richness margalefs index was used as a simple measure of.
The shannonwiener diversity index is a heterogeneity measure that incorporates species richness and evenness hollenbeck and ripple, 2007. Sample values s 60,10,25,1,4 number of species n 5. This presents the shannonwiener also incorrectly known as shannonweaver diversity index for. Shannon weiner index the shannon index is affected by both the number of species and their equitability, or evenness. S total species p i percentage of that species out of the total ln natural log. Species diversity considers both the number of species richness in the community and their relative abundance evenness or equability. The proportion of species i relative to the total number of species p i is calculated, and then multiplied by the natural logarithm of this proportion lnp i. An example of the shannonwiener diversity index for science students. Pdf shannonwiener diversity index has been used extensively in.
Statistics shannon wiener diversity index tutorialspoint. Shannons index of diversity h is derived from information theory, originally in the context of information in telephone systems shannon, 1948. Shannon wiener species diversity index formula data analysis. Hills n2 diversity index 17 s i pi d n 1 2 2 1 1 2 in this formula n2 is the number of numerous species d 1, pi the relative frequency of i species in community. Shannonwiener diversity function h pi lnpi h shannonwiener index of species diversity s number of. It measures the rarity and commonness of species in a community. Sum i individual species e which is the evenness of those species. Shannonwiener diversity index is a term in biologyecology. The shannonwiener diversity index is one measure that we will use to try to draw information from samples in the field. The simpson index is a dominance index because it gives more weight to common or dominant species.
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